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1 Tree was published in this study

Molecular phylogeny of the tribe Sphodrini (Coleoptera: Carabidae) based on mitochondrial and nuclear markers.
Ruiz, Jordal, Serrano. (2009). Mol Phylogenet Evol.
Calathidius accuminatus Licinopsis obliterata Laemostenus terricola Laemostenus complanatus Anchomenidius astur Pristosia aeneola Lindrothius caucasicus Lindrothius pseudopraestans Calathus fuscipes Calathus rotundicollis Calathus mollis Calathus circumseptus Dolichus halensis Synuchus nitidus Synuchus vivalis Synuchus cycloderus Paraeutrichopus harpaloides Paraeutrichopus pecoudi Gomerina calathiformis Platyderus incertans Platyderus varians Amaroschema gaudini Anchomenus dorsalis Agonum muelleri Paranchus albipes Olisthopus fuscatus Poecilus purpurascens Poecilus quadricollis Percus politus Amara aenea Zabrus ignavus Pterostichus globosus Orthomus velocissimus Harpalus rufipes Harpalus affinis Stenolophus teutonus
Tree is not drawn to scale due to some nodes not having time estimates denotes timed nodes (click a timed node to view time)
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