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Bucorvus leadbeateri Bucorvus abyssinicus Rhyticeros plicatus Rhyticeros undulatus Rhyticeros subruficollis Aceros cassidix Aceros nipalensis Penelopides panini Penelopides manillae Aceros corrugatus Aceros corrugatus rugosus Berenicornis comatus Tropicranus albocristatus Bycanistes subcylindricus Bycanistes brevis Bycanistes bucinator Ceratogymna atrata Ocyceros gingalensis Anthracoceros convexus Anthracoceros albirostris Anthracoceros malayanus Anorrhinus austeni Anorrhinus tickelli Anorrhinus galeritus Rhinoplax vigil Buceros rhinoceros Buceros rhinoceros silvestris Buceros bicornis Tockus fasciatus fasciatus Tockus alboterminatus Tockus nasutus Tockus jacksoni Tockus deckeni Tockus leucomelas Tockus erythrorhynchus kempi Tockus ruahae Tockus erythrorhynchus
Tree is not drawn to scale due to some nodes not having time estimates denotes timed nodes (click a timed node to view time)
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