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1 Tree was published in this study

An old adaptive radiation of forest dung beetles in Madagascar.
Wirta, Orsini, Hanski. (2008). Mol Phylogenet Evol.
Paracopris punctulatus Microcopris reflexus Onthophagus signatus Phalops wittei Digitonthophagus gazella Onthophagus depressus Onthophagus hinnulus Onthophagus variegatus Onthophagus flavimargo Onthophagus rectecornutus Proagoderus sappharinus Liatongus vertagus Onthophagus avocetta Tragiscus dimidiatus Drepanocerus laticollis Euoniticellus triangulatus Helictopleurus giganteus Helictopleurus sinuaticornis Helictopleurus corruscus Helictopleurus marsyas Helictopleurus nicollei Helictopleurus carbonarius Helictopleurus minutus Helictopleurus viridiflavus Helictopleurus neoamplicollis Helictopleurus semivirens Helictopleurus steineri Helictopleurus dorbignyi Helictopleurus fungicola Helictopleurus cribricollis Helictopleurus rudicollis Helictopleurus neuter Helictopleurus fasciolatus Helictopleurus fissicollis Helictopleurus splendidicollis Helictopleurus fulgens Helictopleurus politicollis Helictopleurus perrieri Helictopleurus quadripunctatus Helictopleurus unifasciatus Heterosyphus sicardi Oniticellus planatus Onthophagus elegans Parascatonomus penicillatus Tiniocellus spinipes Serrophorus seniculus Liatongus militaris Onitis subopacus
Tree is not drawn to scale due to some nodes not having time estimates denotes timed nodes (click a timed node to view time)
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